Glycogenoses and Glycogen Storage Diseases, General Aspects
General Summary of glycogenosis Classificaiton of glycogenoses by enzyme deficiency
Summary: Glycogenoses are resulted from abnormal metabolism of glycogen or the lack of a particular lysosomal enzyme to degrade glycogen or catabolism of glucose. Most but not all subtypes are associated with excessive glycogen storage. Glycogen storage diseases (GSD) should be reserved to the subtypes with substantial glycogen storage. Clinical manifestations of skeletal muscle inclue weakness with hypotonia, exercise intolerance or muscle cramps with or without myoglobinuria. The liver and the heart are also affected in many subtypes. The muscle biopsy can show striking glycogen storage or minimal changes. The clinicopathologic features are type specific. When glycogen storage is present in the muscle, the most common pathologic findings is subsarcoplasmic and cytoplasmic vacuoles that contain PAS(+) material.
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Diagram
of glycogen metabolism.
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There are 11 types of glycogenosis as classified by
their enzymatic deficiency. Nine of them affect the muscle. Type I (Von Gierke
disease) and VI (Hers disease) do not affect the muscle.
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Many of them have several subtypes with differenc
clinical features. The more severe form of type I (von Gierke), type II (Pompe’s
disease), type IV (Andersen’s disease), and type VI (Tarui’s disease)
glycogenoses are not compatible with prolonged survival. The less severe forms
of these diseases are compatible with long-term survival.
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Some enzymes that are involved in glycogen
metabolism exist in a single isoform (e.g. acid maltase) and its deficiency is
likely to involve many different tissues. Some enzymes exist as several tissue-speficif
isoforms (e.g., phosphroylase) and the tissue being affected as a result of the
deficiency of a particular isoform is tissue specific. Different isoforms may
also be present in mature and immature msuscle.
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Clinical presentation can be weakness and hypotonia
(GSD type II and III), exercise intolerance or muscle cramps with or without
myoglobinuria (GSD type V, VII, VIII, IX, X, XI) or minimal musclar symptoms but
with significant systemic symptoms (GSD type V).
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Depending on the subtype,
the amount of glycogen can vary from dramatic to modest. The form of storage
material may occur in form of PAS(+)-diastase sensitive, PAS(+)-diastase
resistant, and polyglucosan bodies.
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The morphologic changes
are related to the time of biopsy. Muscle biopsy that are obtained after
sternous exercise often reveal necrotic fibers and regenerating fibers.
Summary
of characteristics of glycogenoses:
Head
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Type |
Enzyme |
Gene location |
Clinical
features |
Tissue/system
affected |
Pathology |
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Characterized by weakness &
hypotonia |
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II
(infantile form is also known as Pompe’s disease) |
a-1,4-glucosidase
(acid maltase): this is a lysosomal enzyme |
17(q21-23) |
Severe form: Always fatal in
the first few years of live. Muscle weakness and hypotonia resembles
infantile spinal muscular atrophy (SMA), cardiomegaly and hepatomegaly,
characteristic EKG. Mild form: Resembles
limb-girdle dystrophy |
Muscle,
heart, nervous system, leukocytes, liver, kidney |
Vacuolization,
PAS(+) material in fibers. |
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Amylo-1,6-glucosidase
(debranching enzyme) |
1p21.
Autosomal recessive. |
Mild
weakness and infantile hypotonia. Hypoglycemia and ketosis. Compatible
with long- term survival. |
Muscle,
heart, liver, leukocytes. |
Vacuolization,
PAS(+) material in fibers. |
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Exercise intolerance, fatigue, cramps, myoglobinuria (EIFCM) |
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Glycogen
phosphorylase, M- isoform |
11q13 |
EIFCM.
Other than the fatal infantile form, it is compatible with long-term
survival. |
Entirely
limited to muscle. |
Minimal
or no abnormal storage of glycogen. Many
necrotic and regenerating fibers. |
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Phosphofructokinase-
M isoform |
1(cenàq32) |
EIFCM |
Muscle,
mild hemolytic anemia. |
Minimal
to moderate amount of glycogen accumulation. Polyglucosan bodies. |
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Phosphorylase b
kinase, a-
and b-subunit |
Xq12-q13,
16q12-q13 |
Exercise
intolerance, muscle stiffness, weakness |
Muscle,
liver and heart |
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IX |
Phosphoglycerate
kinase, A-isoform |
Xq13 |
EIFCM |
Muscle,
hemolytic anemia, central nervous system |
Accumulation
of glycogen granules may be light. Biopsy may be histochemically and
histologically normal |
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X |
Phosphoglycerate
kinase, M-mutase |
7 |
EIFCM |
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Accumulation
of PAS(+) material and glycogen content on biochemical analysis. |
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XI |
Lactate
dehydrogenase, M-isoform |
11 |
EIFCM |
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No or minimal muscle symptoms |
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IV
(Andersen’s disease)
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a-1,4-glucan:a1,4-glucan
6-glycosyl transferase (“branching enzyme”; amylo (1,4 à1,6
trans-glucosidase) |
3 |
No
muscle symptoms or mild wasting and weakness |
Muscle,
hepatomegaly, cirrhosis, liver failure, Heart. Death
usually occurs by four years of age because of cirrhosis. |
PAS(+),
PASD(-) material and polyglucosan bodies. |
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VI
(Hers disease)
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Glycogen phosphorylase-
B, M, H (Brain, muscle and liver isoforms) [Chang
S et al., Hum Mol Genet 1998 May;7(5):865-70] |
X-linked form in
Xp22.1–22.2;
autosomal recessive form probably related to glycogen phosphorylase-H
isoform on 14q21–22 |
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Clinical
presentation is heterogenous. Muscle
is not affected. |
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I
(von Gierke disease)
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A
type: glucose-6-phosphatase (G6P)
gene Non-a
type:
glucose-6-phosphate
translocase (G6PT) gene. [Jenecke
et al., 2001 |
G6P is on
chromosome 17; G6PT is on chromosome 11 |
Infantile
onset, often associated with mild mental retardation. Compatible with
long-term survival. Fasting hypoglycemia, fasting lactate acidosis, hepatomegaly, growth retardation, hyperlipidemia, and hyperuricemia. Some patients are prone to severe infections because of neutropenia. Other
system problems include renal failure and osteoporosis. |
Muscle
not affected. The liver,
central nervous system, and other system. |
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Classification
of glycogenoses by deficiency of functions:
Head
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Disease |
Enzyme |
Function |
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Glycose-6-phosphate to glycogen (synthesis of glycogen) |
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a-1,4-glucan:a1,4-glucan 6-glycosyl transferase |
Branching
enzyme |
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I
(von Gierke’s disease) |
Glucose-6-phosphatase |
Glucose-6-phosphateàGlucose |
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Glycogen to fructose-6-phosphate (anabolism of glycogen) |
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a-1,4-glucosidase (acid maltase) |
Lysomal
breakdown of glycogen |
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Amylo-1,6-glucosidase |
Debranching
enzyme |
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Glycogen
phosphorylase, M- isoform |
GlycogenàGlucose-1-phosphate
(muscle) |
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VI
(Hers disease) |
Glycogen
phosphorylase, H- isoform |
GlycogenàGlucose-1-phosphate
(liver) |
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Fructose-6-phosphate to lactate (glycolysis and beyond) |
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Phosphofructokinase-
M isoform |
Fructose-6-phosphateàFructose-1,6-diphosphate |
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IX |
Phosphoglycerate
kinase, A-isoform |
3-P-glycerol
phosphateà 3-phosphoglycerate |
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X |
Phosphoglycerate
mutase, M-subunit |
3-phosphoglycerateà2-phosphoglycerate |
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XI |
Lactate
dehydrogenase, M-isoform |
PyruvateàLactate |
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Control |
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Phosphorylase
b kinase, a- and b-subunit |
Activation
of phosphorylase-b into phosphorylase-a |
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